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Toco Toucan

Ramphastos toco

A blood-vessel-packed bill that doubles as a radiator, and a throat that takes fruit without the tongue ever touching it.

Danger tierLowEffectively harmless to people.
IUCN status [LC]Least ConcernPopulation stable and widespread in natural habitats.Source: IUCN Red List
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A Bill That Sheds Heat

The toco toucan carries the largest beak relative to body size of any bird, and it also works as a radiator. Adult toco toucans can widen the blood vessels in their beaks depending on the temperature, letting heat escape from the bill. At air temperatures above 28°C (82°F), the bill carries away an average of 60 percent of the bird’s non-evaporative heat loss.

A vascular radiator Relative to its size, the toucan’s bill is one of the largest thermal windows in the animal kingdom, rivaling an elephant’s ears in its ability to radiate body heat.

The Biggest Bill for Its Size

The toco is the largest and heaviest toucan, up to 61 centimeters (24 inches) long and 860 grams (1.9 pounds). The oldest toco toucan in the AnAge longevity database lived 16.2 years, in captivity. Its bill can make up 30 to 50 percent of its total body surface area, yet it accounts for just one-twentieth of its mass. Under the surface is a sandwich: a shell of overlapping keratin tiles, each about 50 microns across, around a core of bony foam. The bill is the toucan’s main foraging tool, letting it reach into tree holes and grasp fruit from surrounding branches.

Fruit Thrown to the Throat

Many birds move food to the throat with the tongue. A toco toucan does not. It places a piece of fruit at the very tip of its bill and leans its head back, projecting the food straight into its pharynx. Its tongue never touches the fruit. Tongue movement instead goes with a widening of the throat that lets the food reach the esophagus.

Life in Semi-Open Country

Toco toucans range from Guyana south to Argentina and from central Brazil west to southeastern Peru. The toco is the only non-forest toucan, found mostly in dry, semi-open country with scattered trees, such as woodland and savanna, and along forest edges. In Brazil it is abundant in the cerrado. It relies on seasonal fruit, so it moves from one habitat and region to the next to feed. It eats a great many figs and other fruit, and occasionally insects and other birds’ eggs, including those of hyacinth macaws. When foraging, toco toucans travel in small groups through the canopy, gliding in single file from site to site. Jaguars, snakes, coatis, and eagles are among their predators. The IUCN Red List classifies the species as Least Concern.

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Traits

Intel
& FAQ

01How does the toco toucan's bill act like a radiator?

The bill is heavily supplied with blood vessels, and the bird controls the flow through them by narrowing or widening them. With the vessels widened, the bill gives off heat. Above 28 degrees Celsius it accounts for an average of 60 percent of the toucan's non-evaporative heat loss.

Source: van de Ven et al. 2016

02What is inside a toco toucan's bill?

Foam, of a sort. Materials scientists who compared toucan and hornbill beaks describe the toco's bill as a sandwich: layers of keratin scales outside, and a core that is a fibrous network of bony closed-cell foam. Modelling that design, they found the foam core raises the bill's resistance to bending three to six times while lowering its compressive strength by only 50 percent.

Source: Seki, Bodde & Meyers 2010

03Does a toco toucan really throw its food down its own throat?

In effect, yes. Researchers who studied toco toucans feeding found that the food is projected between the jaws and travels from the tip of the bill to the pharynx in one transport cycle, which they define as ballistic transport. The tongue plays no part in pushing it along and never makes contact with the food.

Source: Baussart et al. 2009

04Does a toco toucan really eat hyacinth macaw eggs?

Occasionally. The toco toucan eats mainly fruit but is an opportunistic feeder, and the eggs it sometimes takes include those of hyacinth macaws. The macaw depends on it all the same: hyacinth macaws almost always nest in manduvi tree hollows, and the manduvi tree relies on toco toucans to disperse its seeds.

Source: Animal Diversity Web, University of Michigan

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