What the sources say
Ocean Sunfish (Mola Mola)Mola mola
Every figure, record and mechanism on the Ocean Sunfish (Mola Mola) profile was written from a source, and this is the working table behind it: 73 claims checked word for word against these sources. The left column is what the page says. The right column is the source's own words, copied, not paraphrased; a ... marks a cut and[brackets] an insertion. Open any source and check it yourself.
Profile updated
Source
01
Animal Diversity Web, University of Michigan
Mola mola (Ocean Sunfish). Animal Diversity Web, University of Michigan.
- The page saysLarge sharks often prey on sunfish
- The source says
Ocean sunfish are often preyed upon by large sharks and California sea lions
Note: ADW is dated 2011; its sea-lion half is narrowed by FishBase's current page (next claim) - The page sayscleaner fish such as wrasses and angelfish pick off its parasites when it occasionally visits kelp beds and reefs
- The source says
occasionally venture into kelp beds and deep coral reefs in order to be cleaned of parasites by fishes such as wrasses (Labroides dimidiatus and Thalasoma lunare) and Emperor Angelfish
- The page saysThe clavus is the rudder-like structure at the rear of the fish
- The source says
they instead have a clavus, which is a truncated tail, used more like a rudder than for propulsion
Source
02
FishBase
Mola mola (Linnaeus, 1758), Ocean sunfish. FishBase.
- The page saysUp to 1.3 tonnes
- The source says
max. published weight: 1.3 t
Note: 1.3 t worded as 1.3 tonnes - The page saysthen floats flat at the surface between dives
- The source says
Individuals often drift at the surface while lying on its side
- The page saysTemperate and tropical oceans worldwide
- The source says
Circumglobal: tropical and temperate zones of all oceans.
Note: backs stats.habitat - The page saysA body with no tail fin, steered by a stiff clavus rudder
- The source says
The caudal fin is replaced by a rudder-like structure called
Note: backs stats.superpower - The page saysThe ocean sunfish (Mola mola) can grow past 3 meters long and weigh up to 1.3 tonnes
- The source says
Max length : 333 cm TL male/unsexed; ... max. published weight: 1.3 t
Note: 333 cm TL is worded as 'past 3 meters'; a maximum, so 'can grow' - The page saysCalifornia sea lions go after the juveniles in Monterey Bay
- The source says
Juveniles are victims of California sea lions in Monterey Bay
Note: 'go after' is the phrasal verb for hunting (the source's 'are victims of'), not a sequence; FishBase is current and narrows ADW 2011's general sea-lion line - The page saysA Mola mola can grow past 3 meters long and weigh up to 1.3 tonnes
- The source says
Max length : 333 cm TL male/unsexed; ... max. published weight: 1.3 t
Note: 333 cm TL is worded as 'past 3 meters' - The page saysIts name comes from the Latin for millstone
- The source says
Etymology: Mola: Latin, mola, -ae = stone mill; because of the shape of this fish
- The page saysOne 1.5-meter female was found carrying about 300 million eggs, the largest clutch estimate for the species
- The source says
300 million eggs found in a 1.5 m long female ... This is the largest clutch estimate for this species
Note: was 'A female can carry roughly 300 million eggs in one spawning': the figure is one 1.5 m female, and FishBase calls the species a multiple spawner, so not one spawning
Source
03
Oceana
Ocean Sunfish. Oceana.
- The page saysthen floats flat at the surface between dives
- The source says
Because they must sometimes dive into deep, cold waters to catch their prey, these fish will warm up by basking on their side at the water
Note: the 'between dives' half - The page saysjuveniles can pack on up to 0.82 kilograms a day
- The source says
they can grow up to 0.82 kilograms (1.8 pounds) every single day
- The page saysconsistent with basking between dives to recover warmth
- The source says
these fish will warm up by basking on their side at the water
Note: backs 'basking' - The page saysThey are most likely warming back up after cold, deep dives
- The source says
Because they must sometimes dive into deep, cold waters to catch their prey, these fish will warm up by basking on their side at the water
Note: hedged 'most likely' because Nakamura et al. 2015 only 'suggest' it and the Australian Museum quotes Potter and Howell 2010 finding no evidence for it; reviewer cut 'not sunbathing for fun'
Source
04
Australian Museum
Ocean Sunfish, Mola mola. Australian Museum.
- The page sayswith no tail fin
- The source says
These interesting fish are completely without a caudal fin
Note: reviewer: replaces 'finless'; sunfish have dorsal, anal and pectoral fins, only the caudal fin is missing - The page saystagged fish spent most of their time in the top 200 meters
- The source says
over 80% of time in the top 200 m
Note: backs stats.habitat; kept at the tagging study's scope; the Australian Museum page quotes this figure from Potter and Howell 2010 - The page saysand one dived to 844 meters
- The source says
The maximum depth recorded by any fish was 844 m.
Note: backs stats.habitat; the page quotes this from Potter and Howell 2010. Reviewer cut 'to feed', which no source ties to that dive - The page saysJellyfish, comb jellies and siphonophores
- The source says
Ocean Sunfish feed on jellyfish, salps, ctenophores
Note: backs stats.diet; 'comb jellies' is the common name for ctenophores; siphonophores backed by nakamura-2015 below. Reviewer dropped 'as adults': no source gives the adult diet as such - The page saysthe clavus, the sunfish's rudder-like rear end, has a wavy, frilled edge in Mola mola and a rounded one with no indent in M. alexandrini
- The source says
Mola mola can be identified from its wavy (frilly) clavus edge, 11-14 clavus fin rays ... (the clavus is round without an indent in Mola alexandrini and rounded with an indent in Mola tecta)
Note: reviewer: 'smooth' replaced by the source's own 'without an indent' - The page saysSunfish have no tail fin
- The source says
These interesting fish are completely without a caudal fin
- The page saysone tracking study logged a dive to 844 meters
- The source says
The maximum depth recorded by any fish was 844 m.
Note: the page quotes this from Potter and Howell 2010 - The page saysa nod to its rounded shape
- The source says
the name Mola comes from the Latin word for millstone and refers to the rounded shape of the fish
Note: was 'round, flattened shape'; neither source says flattened - The page saysits family, Molidae, includes the largest bony fish in the world
- The source says
Sunfish are from the family Molidae and are the largest bony fish in the world.
Source
05
Jing et al. 2015
Jing et al. (2015). Mola mola. The IUCN Red List of Threatened Species 2015: e.T190422A1951231.
- The page saysVulnerable
- The source says
IUCN Red List category: Vulnerable (VU).
Note: backs stats.conservationStatus; FishBase lists the same category under Red List version 2025-2 (assessed 07 June 2011), so it is current
Source
06
Sawai & Nyegaard 2022
Sawai & Nyegaard (2022). A review of giants: Examining the species identities of the world's heaviest extant bony fishes (ocean sunfishes, family Molidae). Journal of Fish Biology.
- The page saysFor a long time it also carried a bigger title: the world's heaviest bony fish, at around 2,300 kilograms
- The source says
The world's heaviest extant bony fish has long been considered the ocean sunfish Mola mola (family Molidae). However, recent taxonomic research suggests the world record specimen (2300 kg) was actually a misidentified giant sunfish M. alexandrini.
Note: 'for a long time' for the source's 'long been considered' (was 'for decades') - The page saysA 2022 review of giant-sunfish records found that the specimen behind that number wasn't Mola mola
- The source says
we confirm that the world's heaviest extant bony fish specimen actually weighed was a female M. alexandrini (2300 kg, 272 cm total length) captured from Kamogawa, Chiba Prefecture, Japan, on August 16, 1996.
Note: 2022 is the paper's own publication year; 'wasn't Mola mola' reflects the paper's own confirmed finding, not the earlier hedged 'suggests' sentence - The page saysIt was a giant sunfish, Mola alexandrini, a separate species caught off Japan in 1996
- The source says
the world's heaviest extant bony fish specimen actually weighed was a female M. alexandrini (2300 kg, 272 cm total length) captured from Kamogawa, Chiba Prefecture, Japan, on August 16, 1996
- The page saysthe same review concluded M. alexandrini is the only living bony fish researchers could confirm above 2,000 kilograms
- The source says
we conclude that M. alexandrini is the only extant bony fish species we could confirm that exceeds 2000 kg
Note: 'living' restores the source's 'extant'; 'above' is the source's 'exceeds' - The page saysSo Mola mola is enormous, just not the outright record-holder its old reputation claims
- The source says
we confirm that the world's heaviest extant bony fish specimen actually weighed was a female M. alexandrini
Note: follows from the confirmed record specimen being M. alexandrini; 'enormous' rests on the 1.3 t above - The page saysNo, though it was long thought to be
- The source says
The world's heaviest extant bony fish has long been considered the ocean sunfish Mola mola
Note: was 'Not anymore', which implied the fish once held the record; the record fish was never Mola mola - The page saysThe title rested on a specimen weighed at about 2,300 kilograms
- The source says
the world record specimen (2300 kg) was actually a misidentified giant sunfish M. alexandrini
- The page saysa 2022 review of giant-sunfish records found that fish had been misidentified
- The source says
we confirm that the world's heaviest extant bony fish specimen actually weighed was a female M. alexandrini (2300 kg, 272 cm total length) captured from Kamogawa, Chiba Prefecture, Japan, on August 16, 1996.
Note: 2022 is the paper's own publication year; reflects the paper's own confirmed finding, not the earlier hedged 'suggests' sentence - The page saysit was actually a giant sunfish, Mola alexandrini, a separate species caught off Japan in 1996
- The source says
captured from Kamogawa, Chiba Prefecture, Japan, on August 16, 1996
- The page saysThe review concluded Mola alexandrini is the only living bony fish researchers could confirm above 2,000 kilograms
- The source says
we conclude that M. alexandrini is the only extant bony fish species we could confirm that exceeds 2000 kg
Note: 'living' restores the source's 'extant'; 'above' is the source's 'exceeds' - The page saysso the true ocean sunfish is a very big fish, just not the outright record-holder its old reputation claims
- The source says
we confirm that the world's heaviest extant bony fish specimen actually weighed was a female M. alexandrini
Note: follows from the confirmed record specimen being M. alexandrini
Source
07
Watanabe & Sato 2008
Watanabe & Sato (2008). Functional Dorsoventral Symmetry in Relation to Lift-Based Swimming in the Ocean Sunfish Mola mola. PLoS ONE.
- The page sayscartilage-and-jelly fish
- The source says
they are neutrally buoyant independent of depth because of their subcutaneous gelatinous tissue
Note: the 'jelly' half - The page saysdriven by synchronized, penguin-style dorsal and anal fin strokes
- The source says
they stroked their dorsal and anal fins synchronously ... to generate a lift-based thrust, as penguins do using two symmetrical flippers
Note: backs stats.superpower - The page saysThe dorsal and anal fins do the work instead, beating side to side in sync
- The source says
Lateral acceleration oscillated regularly, showing that the fish stroked their dorsal and anal fins from side-to-side
- The page saysthe way a penguin flaps both flippers together, to generate lift-based thrust rather than a drag-based push
- The source says
demonstrating that the thrust was lift-based rather than drag-based. This pattern of acceleration was previously reported in penguins [7] , which flap both flippers synchronously, indicating that ocean sunfish oscillate the dorsal and anal fins synchronously
- The page saysTagged sunfish cruised at 0.4 to 0.7 meters per second, in the same range as salmon and marlin
- The source says
sunfish indeed swam continuously with frequent vertical movements at speeds of 0.4-0.7 m s-1, which is similar to the records of other large fishes such as salmons, marlins, and pelagic sharks
Note: kept at the tagged fish's scope; was 'That gets a sunfish cruising', which made the fin stroke the cause of the measured speed - The page saysThis is not the limp drifter its reputation suggests
- The source says
Contrary to the common view that ocean sunfish are planktonic fish at the mercy of oceanic currents, the sunfish tagged in this study swam actively
- The page saysa thick gelatinous layer under the skin made up 44 percent of the weight of one 247-kilogram fish
- The source says
The proportion of the mass of the gelatinous tissue to body mass significantly increased with body size ... to 44% for a 247-kg individual
Note: was 'can make up nearly half a large fish's weight', from one 247-kg individual - The page sayskeeping the whole fish neutrally buoyant without a swim bladder
- The source says
sunfish lack a swimbladder, we found that they are neutrally buoyant independent of depth because of their subcutaneous gelatinous tissue that has low density and is incompressible
- The page saysActual propulsion comes from those same dorsal and anal fins beating side to side in unison, generating lift-based thrust the way a penguin's two flippers do
- The source says
they stroked their dorsal and anal fins synchronously ... to generate a lift-based thrust, as penguins do using two symmetrical flippers
- The page saysfor a fish with no tail fin that swims by stroking its dorsal and anal fins side to side in sync
- The source says
they stroked their dorsal and anal fins synchronously ... to generate a lift-based thrust, as penguins do using two symmetrical flippers
Note: task #812, vibe.reason; 'no tail fin' is claimed elsewhere from australian-museum-mola-mola
Source
08
Sims et al. 2009
Sims et al. (2009). Long-Term GPS Tracking of Ocean Sunfish Mola mola Offers a New Direction in Fish Monitoring. PLoS ONE.
- The page saysone GPS-tagged fish covered an estimated 1,819 kilometers over 92 days
- The source says
Sunfish 3, the largest of the fish tagged (1.0 m TL), was tracked for 92 days ... This fish covered an estimated distance of 1,819 km
Source
09
Nakamura, Goto & Sato 2015
Nakamura, Goto & Sato (2015). Ocean sunfish rewarm at the surface after deep excursions to forage for siphonophores. Journal of Animal Ecology.
- The page saysIt dives for siphonophores
- The source says
Siphonophores were the most abundant prey items captured by ocean sunfish
- The page saysJellyfish, comb jellies and siphonophores
- The source says
Siphonophores were the most abundant prey items captured by ocean sunfish
Note: backs the 'siphonophores' part of stats.diet - The page saysCamera tags have caught sunfish hunting siphonophores on repeated daytime dives to roughly 100 to 200 meters, in water colder than 12°C
- The source says
Siphonophores were the most abundant prey items captured by ocean sunfish and were typically located at a depth of 50-200 m where the water temperature was <12 °C ... foraged mainly at 100-200 m depths during the day
Note: 'colder than 12°C' is worded from the source's '<12 °C'; was 'caught adults', but the abstract gives no size or age for the tagged fish - The page saysDeep dives cost heat
- The source says
The body temperatures decreased during deep excursions
Note: was 'Every dive costs heat'; the source says deep excursions - The page saysA sunfish's body temperature drops on the way down
- The source says
The body temperatures decreased during deep excursions
- The page saysresearchers found the rate of heat exchange runs 3 to 7 times greater at the surface than while cooling
- The source says
the whole-body heat-transfer coefficient between sunfish and the surrounding water during warming was 3-7 times greater than that during cooling
Note: 'rate of heat exchange' is plain-English for the source's 'whole-body heat-transfer coefficient'; warming happens 'during subsequent surfacing periods' - The page saysconsistent with basking between dives to recover warmth
- The source says
these results suggest that the main function of surfacing is the recovery of body temperature
Note: 'consistent with' preserves the source's own hedge ('these results suggest'); reviewer cut 'rather than relax' - The page sayssunfish diving repeatedly to about 100 to 200 meters, into water colder than 12°C, to hunt siphonophores
- The source says
Siphonophores were the most abundant prey items captured by ocean sunfish and were typically located at a depth of 50-200 m where the water temperature was <12 °C ... foraged mainly at 100-200 m depths during the day
Note: 'colder than 12°C' is worded from the source's '<12 °C' - The page saysEach deep dive drops their body temperature
- The source says
The body temperatures decreased during deep excursions
- The page saysresearchers found the rate of heat exchange runs 3 to 7 times greater once the fish is back at the surface than it did while cooling
- The source says
the whole-body heat-transfer coefficient between sunfish and the surrounding water during warming was 3-7 times greater than that during cooling
Note: 'rate of heat exchange' is plain-English for the source's 'whole-body heat-transfer coefficient' - The page saysconsistent with time at the surface being how it rewarms between hunting trips
- The source says
these results suggest that the main function of surfacing is the recovery of body temperature
Note: 'consistent with' preserves the source's own hedge ('these results suggest'); was 'lying flat in the sun', but the source says surfacing and warm surface water
Source
10
Sousa et al. 2016
Sousa et al. (2016). DNA barcoding identifies a cosmopolitan diet in the ocean sunfish. Scientific Reports.
- The page sayssmaller sunfish eat mainly crustaceans and small fish
- The source says
smaller individuals feeding mainly on small crustaceans and teleost fish
Note: backs stats.diet; keeps the source's 'mainly' - The page saysWhat a sunfish eats changes as it grows
- The source says
we found evidence for an ontogenetic shift in the diet, with smaller individuals feeding mainly on small crustaceans and teleost fish, whereas the diet of larger fish included more cnidarian species
- The page saysIn a study of juveniles and sub-adults off southern Portugal, all 37 centimeters to 1.1 meters long
- The source says
Total length of the 100 sampled individuals ranged from 0.37 to 1.10 m, corresponding to juveniles and sub-adults ... sunfish captured in the tuna set-net off southern Portugal
Note: reviewer: puts the diet study at its own scope; it sampled no adults - The page saysthe smaller fish ate mainly small crustaceans and fish from the seabed and coastal waters
- The source says
smaller individuals feeding mainly on small crustaceans and teleost fish ... smaller sunfish feed predominantly on benthic and on coastal pelagic species
Note: 'from the seabed and coastal waters' is the source's 'benthic and ... coastal pelagic species' - The page sayswhile the larger ones relied on open-water prey and ate more kinds of jellyfish and their relatives
- The source says
whereas the diet of larger fish included more cnidarian species ... whereas larger fish depend on pelagic prey
Note: 'jellyfish and their relatives' for cnidarians; 'more kinds' for 'more cnidarian species'. Was 'larger adults ... especially siphonophores': the study sampled no adults, and siphonophores come from a different study
Source
11
Sawai et al. 2017
Sawai et al. (2017). Redescription of the bump-head sunfish Mola alexandrini (Ranzani 1839), senior synonym of Mola ramsayi (Giglioli 1883), with designation of a neotype for Mola mola (Linnaeus 1758) (Tetraodontiformes: Molidae). Ichthyological Research.
- The page saysThese species had long been confused for each other
- The source says
as these two species have long been confused
- The page saysa 2017 taxonomic study designated a fixed reference specimen for Mola mola to sort them out
- The source says
A neotype of M. mola is designated for comparison with M. alexandrini, as these two species have long been confused.
Note: 2017 is the paper's own publication year
Source
12
Johnson & Britz 2004
Johnson & Britz (2004). Leis' conundrum: Homology of the clavus of the ocean sunfishes. 2. Ontogeny of the median fins and axial skeleton of Ranzania laevis (Teleostei, Tetraodontiformes, Molidae). Journal of Morphology.
- The page saysa study tracing its development in a related sunfish found it is formed from modified elements of the dorsal and anal fins, not a leftover tail fin: the caudal fin is lost in the sunfish family
- The source says
the molid clavus is unequivocally formed by modified elements of the dorsal and anal fin and that the caudal fin is lost in molids
Note: the study traced the slender sunfish Ranzania laevis (title) and concluded for the family Molidae
Source
13
Davenport et al. 2018
Davenport et al. (2018). The locomotor system of the ocean sunfish Mola mola (L.): role of gelatinous exoskeleton, horizontal septum, muscles and tendons. Journal of Anatomy.
- The page saysThat gelatinous layer is roughly 90 percent water, threaded with a collagen and elastin meshwork
- The source says
The capsule is gelatinous (89.8% water) with a collagen and elastin meshwork.
Note: 89.8% rounded to 'roughly 90 percent' - The page saysit houses the muscles that drive the dorsal and anal fins
- The source says
It provides two compartments, one for dorsal fin musculature and one for anal fin muscles
Source
14
Pan et al. 2016
Pan et al. (2016). The genome of the largest bony fish, ocean sunfish (Mola mola), provides insights into its fast growth rate. Gigascience.
- The page sayscartilage-and-jelly fish
- The source says
its endoskeleton is mainly composed of cartilage
Note: the 'cartilage' half; the 'jelly' half is the subcutaneous gelatinous tissue, next claim - The page saysA body with no tail fin, steered by a stiff clavus rudder
- The source says
its lack of a caudal fin, which is replaced by a broad and stiff lobe
Note: backs 'stiff' (was 'rigid', which no source used) - The page saysThe skeleton is mostly cartilage
- The source says
its endoskeleton is mainly composed of cartilage
Note: reviewer cut the lead-in 'Most of that bulk isn't muscle or bone', which no source states - The page saysResearchers studying the sunfish genome found several genes tied to growth-hormone signaling showing signs of positive selection or sped-up evolution, which the study says might explain its fast growth rate
- The source says
Several sunfish genes involved in the growth hormone and insulin-like growth factor 1 (GH/IGF1) axis signalling pathway were found to be under positive selection or accelerated evolution, which might explain its fast growth rate and large body size
Note: was 'under unusually strong selection', which the source does not say
How the table was made: a writer drafted the profile from these sources and logged each claim with its quote, a reviewer with nothing but this table checked every sentence said no more than its quote, and a script confirmed each quote appears word for word in the source. The method is in the site'sAbout page.